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Key to species of subgenus Ateuchetus
1. Posterior margin of metafemur not emarginate . .. .. .... .......... .... .. ........ ...... 2
-- Posterior margin of metafemur emarginate . .. ........ ... .. .. .. ... .. .. .... ...... .... .... 3
2.
Pronotum as well as elytra covered with large, shallow and opaque
punctae, elytral striae barely noticeable. Length 11-25 mm. Fig. 1
.. . .... .. .......... ...... ... ..... .. . .. . ... .. . .. . .. ..... .. .. . .... ....... . .... ... S. variolosus
--
Pronotum with small punctae, elytra devoid of punctae. Elytral interstriae
gently convex; elytral striae mesially glossy and laterally opaque, resulting
in appearance of pronounced striation (some populations may have flat
interstriae and inconspicuous striae, but punctation of
pronotum does not
change). Length 11-25 mm ... .......... ................. .... .. ... .............. S. laticollis
3.
Pronotum covered with large, shallow and opaque punctae ....... ... ........ . 4
--
Pronotum with sparse and unevenly distributed fine punctae . .. ......... .. .... 5
4. Elytra with punctae as large, shallow and opaque as
pronotum.
Emargination of metafemur short, confined to
proximal third of posterior
margin. Length 13-27 mm. Fig. 3 ... .. ................ .. .... .. .. S. cicatricosus
-- Elytra with sparse, fine punctae. Emargination of metafemur long,
occupies two-thirds of posterior margin. Length 14-28 mm. Fig. 4
....... .... ... .... ... .. ..... ........ ...... ..... ....... ........... ...... ..... ......... . S. semipunctatus
5. Dorsal mesial carina of
protibia complete throughout length.
Basal
margin of
pronotum oblique (incomplete), with only sparse, fine punctae.
Length 11-25 mm. Fig. 5 .................................. .... .. ... .... .... S. armeniacus
-- Dorsal mesial carina of
protibia complete in
distal third, then briefly
interrupted and proximally replaced by row of granules.
Basal margin of
pronotum sharp (complete), elevated and with row of confluent punctae.
Length 11-25 mm. Fig. 6 .......................................... S. puncticollis
Key to species of subgenus Mesoscarabaeus
1.
Protibia with pronounced denticles at
bases of three
proximal lateral
teeth. ....... .... .. .................... .... ........ ...... ... .... ........ .... .. ... ...... ......... ... ....... .. .. 2
--
Protibia with pronounced denticle only at
base of proximalmost lateral
tooth . ... ......... ................ ....... ..... ..... ..... .... ......... ..... ...... ..... .. .... .... ... ....... ..... 3
2. Head
vertex with conspicuous impression. Mesial margin of
protibia
sparsely and coarsely serrate. Length 20-34 mm. Fig. 7 .. ... S. andrewesi
-- Head
vertex without impression. Mesial margin of
protibia densely and
finely serrate. Length 22-30 mm. Fig. 8 .... ...... .. .... ... .. .... .. .. S. bannuensis
3. Head with frontal tubercle strong, sharp and well pronounced forward.
Head
vertex with conspicuous impreSSion. Ventral margin of
clypeus with
sharp sagittal tooth. Profemur with
apical tooth on dorsal side. Length 18-
31 mm. Fig. 9 .................................. .... . ...... ........... ...... S. cristatus
-- Head with frontal tubercle weak, blunt and only slightly pronounced
forward. Head
vertex without impreSSion. Ventral margin of
clypeus
without sagittal tooth. Profemur without
apical tooth on dorsal side.
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 4
4. Punctae of
pronotal disk smaller than largest granules near posterolateral
angles. Impunctate central track of
pronotum narrow, with poorly defined
borders. Anterior termination of metasternal bulge between mesocoxae
inclined forward. Asian species occurring in India, Pakistan and s. Iran.
Length 19-23 mm. Fig. 10 ...... ........ .... ........... ..... .......... ....... S. brahminus
-- Punctae of
pronotal disk markedly larger than largest granules near
posterolateral angles. Impunctate central track of
pronotum wide, with
well defined borders. Anterior termination of metasternal bulge between
mesocoxae vertical. Afrotropical species extending into Palearctic region
only in Egypt. Length 19-25 mm. Fig. 11 .. . ... ...... ...... ... S. sennaariensis
Key to species of subgenus Scarabaeus s. str.
1. Eyes small, their dorsal width about equal to one-third of posterior genal
margin. Crenate
basal margin of
pronotum not separated from remaining
surface by furrow. Frontoclypeal carina weak, mesially interrupted,
without tubercles ....... .... ......... .... .... ...... ........ .... ....... .... .. ...... ....... .... ... .... .. 2
-- Eyes large, their dorsal width about equal to one-half of posterior genal
margin. Crenate
basal margin of
pronotum separated from remaining
surface by thin fur. ... .... ... .. ... ...... .. .. .... .... ..... ..... .... .... ... .. ....... .... .. ...... ..... .... 3
2. Emargination between mesial clypeal teeth semicircular, that between
mesial and lateral clypeal teeth nearly triangular.
Pronotum sparsely
covered by fine granules. Elytral interstriae finely punctate. Body opaque.
Length 15-30 mm. Fig. 12 ..... .. .. ... .... .... .. ..... ... .......... .. .. . .. ... .. S. pius
-- Emargination between all clypeal teeth semicircular.
Pronotum densely
covered by coarse granules that mesially and posteriorly change to
dense, coarse, deep punctae. Elytral interstriae with large, shallow,
opaque punctae. Body semi-glossy. Length 20-35 mm. Fig. 13
..... ............ ........... .. ..... .. .. .. ... ..... .... .. ... .... ... ..... ..... .. .. .. .... ..... ........ .. S. wi/son;
3.
Protibia with conspicuous denticles at
bases of two (sometimes three)
distal lateral teeth. Frontoclypeal carina weak, medially interrupted,
without tubercles. Metatibia lacks mesial brush of contiguous setae in
either sex. Length 22-35 mm. Fig. 14 .. ...... .... ....... ......... ... . S. gangeticus
--
Protibia with conspicuous denticle only at
base of
proximal most lateral
tooth. . ....... ....... ..... .... .... .. .. ...... .. ...... ...... ......... ... ... .. .... ... .... ......... .... ... ... ..... 4
4. Weak V-shaped carina on ventral side of third protibial tooth does not
reach anywhere near central longitudinal carina. Body longer, with nearly
parallel elytral margins .. .. ... ... .... ....... . ..... . .... .... .. ... ... .. .. . ..... . ... ....... 5
-- Weak V-shaped carina on ventral side of third protibial tooth reaches or
nearly reaches central longitudinal carina. Body wider, with rounded
elytral margins . ....... ...... .. .... .. ... .... .. .. ........ ..... .... .. ..... ....... .. .... ........ ....... ..... 6
5. Lateral margin of
protibia between
proximal end and proximalmost tooth
widens forward only slightly, causing said tooth to project strongly
laterally. Metatibial brush reddish yellow. Tarsal claws normal. Tips of
parameres sharp and curved down. Body semi-glossy. Length 25-40 mm.
Fig. 15 ..... ......... ... ..... .... ......... ...... ....... .. ..... .... .... ..... ...... .... .... S. acuticollis
-- Lateral margin of
protibia between
proximal end and proximalmost tooth
widens forward markedly, causing said tooth to project only slightly.
Tarsal claws reduced. Tips of parameres dorsoventrally flattened and
curved laterally. Body glossy. Length 25-38 mm. Fig. 16
... ... ... ... ..... ... . ... .. .... .... ......... ........ .... .. ..... .. .. ... ... . S. transcaspicus
at Meso- and metatibial
pubescence dark brown to black.
.... .. .... .... . ..... .... .... ...... .......... .. ....... ... S. transcaspicus transcaspicus
bt Meso- and metatibial
pubescence reddish brown.
.... ... ... .. ..... . ...... ... .... .. ..... .. .... ........ .. ..... S. transcaspicus aegyptiacus
6. Frontoclypeal carina sagittally uninterrupted, with two closely spaced and
sharp mesial tubercles. Body slightly glossy. Length 20-36 mm. Fig.
17 ...... ...... .... ...... .... ....... ... ...... ...... .. ...... ..... ...... ......... ..... .. ..... .... S. carinatus
-- Frontoclypeal carina sagittally interrupted, gap bordered by more or less
developed tubercles . ..... .... .... ... .... .. ........... ... .... .... ..... .. .... .... .... ... ......... ..... 7
7. Emarginations between clypeal teeth mesially semicircular, laterally
angular (excision-like). Body nearly opaque. Length 16-33 mm. Fig. 18
..... ... .... ... ..... ..... ....... .... .. ... .... ...... .... .......... .. .. ... .. .. .... .. ... ....... ........ S. typhon
-- All emarginations between clypeal teeth semicircular . ...... .. ............. .... ... 8
8. Entire
pronotum granulose and punctate, without notable smooth spots.
Dorsal mesial longitudinal carina of
protibia proximally composed of fine
granules . .. ..... .... .. .... ... ..... .... .. .... ..... ....... ... .......... ..... .... .. .. .... ... .. ....... ...... .. . 9
--
Pronotal sculpture interrupted by notable smooth spots. Dorsal mesial
longitudinal carina of
protibia proximally composed of coarse granules.
....... ...... .... ......... .... ....... ...... ...... .................................... ....... ..... ... ... .......... 1 0
9.
Pronotal disk with fine punctae scattered between large, primary
punctae. Paramere terminations broad-based, with sharp and laterally
curved tips. Length 20-40 mm. Fig. 19 ....................................... S. babori
--
Pronotal disk with all punctae of equal size. Paramere terminations not
broad-based, with sharp tips curved down and only slightly out. Length
27 -34 mm. Fig. 20 .. .. .... ...... .. .. ................................................. S. winkleri
10. Metatibial brush absent in both sexes. Tips of parameres truncate and
not curved down. Species endemic to Socotra Island. Length 30-35 mm.
Fig. 21 ......................................................... ................ .. .. .. S. dioscoridis
-- Species not occurring in Socotra Island ....................... .. ........ .. .. .. .. .. ...... 11
11.
Pronotal disk finely and sparsely punctate, smooth areas in front of
pronotal base large and conspicuous. Body glossy. Length 29-36 mm.
Fig. 22 ... ... ... ..... ... . ... ..... . ... ... ... .... .. ... ... ... ............... S. zagrosensis
--
Pronotal disk punctate and granulose, smooth areas in front of
pronotal
base small and not conspicuous. Species with silky gloss.
. .. . .. ... . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 1 2
12. Terminal lobe of mesotibia markedly shorter than spur. Metatibia with
mesial brush of contiguous setae only in males. Length 20-40 mm. Fig.
23 ........................ .. ................................................................. .... .. S. sacer
-- Terminal lobe of mesotibia about as long as spur. Metatibia with mesial
brush of contiguous setae present in both sexes. Length 30-40 mm. Fig.
24 .......................................................................... ... ..... .... ... .. S. irakensis
merci pour ta publi !!!! :-) là j'avoue que c'est un peu compliqué avec ces photos... 
